Social insects
"What is it that governs here? What is it that issues orders, foresees the future, elaborates plans and preserves equilibrium?"
Maurice Maeterlinck, The Life of the White Ant, London: George Allen and Unwin, 1927

"Go to the ant, thou sluggard; consider her ways and be wise"
The Bible, Proverbs, 6.6

Who is "in charge" of an ant colony?
  Stigmergy occurs when one insect modifies the environment and the other insect reacts to the modified environment
E.g. chemical trails, response of termites to nest structure.
 
 
Social insects  Swarm intelligence applications
Ant foraging behaviour using indirect recruitment optimization/routing in communications networks
e.g. travelling salesman problem
Division of labour/task allocation differentiation in multiagent system or group of robots
Cooperative prey transport cooperative transport in robotics
 
Ref. Eric Bonabeau, Marco Dorigo and Guy Theraulaz, Swarm Intelligence: from Natural to Artificial Systems, New York/Oxford: Oxford University Press, 1999.